Memphis Depay's second-half introduction sparked a remarkable fightback that saw Moussa Dembele and Leo Dubois net 119 seconds apart at an empty Groupama Stadium.

Lyon, shock 2-1 winner at Manchester City a fortnight ago, was forced to play this match beyond closed doors as punishment for crowd disturbances during last season's Europa League, and the eerie atmosphere initially played into Shakhtar's hands.

Junior Moraes struck a double either side of half-time to put the visitor within reach of a second Champions League away victory in 10 attempts but, despite its early dominance, Dembele and Dubois' rescue act proved enough to keep the Ligue 1 club top of Group F.