Rennes pulled off the biggest surprise of Sunday's (AEST) Ligue 1 fixtures with a 2-1 win at title-hopeful Lyon, while Bastia and Angers continued their strong starts.

Pedro Henrique gave the visitor an early lead before Nabil Fekir restored parity for Lyon but Mehdi Zeffane came back to haunt his former club by restoring Rennes' advantage on 55 minutes and it held on for the win.

Mathieu Valbuena, Jordan Ferri and Maxwell Cornet all missed chances for Lyon in a frantic finale.

The home fans made their disapproval clear as they booed last season's Ligue 1 top-scorer Alexandre Lacazette when he was replaced on the hour-mark. 

That result elevated Rennes to fifth in the table, one place below unbeaten Nantes which secured a 1-0 win over Reims at the Stade de la Beaujoure, courtesy of a second-half strike from Ermir Lenjani.

Bastia was the day's biggest winner, trouncing struggling Guingamp 3-0 to move up to second, with Julian Palmieri and Lassana Coulibaly finding the net to add to an own goal from Maxime Baca.

Surprise packet Angers round out the top three after an impressive 2-0 victory at GFC Ajaccio, a brace from Cheikh N'Doye proving enough to down the Corsican side.

Those wins see Bastia and Angers level on seven points and plus-four goal difference, with the former ahead on goals scored, with only defending champion Paris Saint-Germain (nine points) above them in the standings.

Monaco remains unbeaten this season after its 1-1 draw at Toulouse, although  it did have to come from behind after Tongo Doumbia gave the host the lead after 21 minutes.

Thomas Lemar earned a point for Leonardo Jardim's men when he came off the bench and fired home a low shot midway through the second half, while in Nice, Caen dropped to sixth after suffering its first loss of the campaign - going down 2-1 to the home team.